During the Evacuation of North Gate, APC-One was crewed by Sgt.Marcus Fenix, Cpl.Dominic Santiago, and Pvt.Federic Rojas. Dom drove the APC at the lead of the convoy, while Rojas manned the gun. After arriving at the North Gate Agricultural Depot, Dom was directed to park APC-One by a group of Sappers. As the convoy prepared to move out, Dom moved APC-One into position at the head of the convoy, and led the convoy out when it was time to move. As they continued on, they began feeling vibrations on the ground, indicating that Locust were about to attack. After reaching the Timgad Bridge, APC-One guarded the bridge while the convoy crossed over it. When Cpl.Damon Baird came under attack while assisting a stalled vehicle, Dom began driving APC-One to assist him. They picked up Baird, along with Sgt.Bernadette Mataki, Pvt.Augustus Cole, and driver Jeff Tatton, and then began heading back to Jacinto. After communication was lost with Col.Victor Hoffman and Pvt.Tai Kaliso, Marcus and Dom left APC-One to rescue them, and Cole drove the vehicle back to Jacinto.